https://www.antweb.org/description.do?genus=harpegnathos&species=saltator&rank=species
From AntWeb.org:
Biology: This species has an unusual colony structure compared to other ponerine genera. In the beginning, young winged queens fly out from their natal colony, mate with foreign males and start new colonies independently. Once the founding queen dies (after a few years), several daughter workers replace her as reproductives in the colony. It is characteristic of several ponerine ants that workers can mate (almost always with foreign males) and store sperm in the spermatheca. In H. saltator, many young workers inbreed with males from the same colony (their brothers). They fight to establish a dominance hierarchy, and a few high-rankers will become reproductives and lay eggs, so-called gamergates. Every year, new winged queens will be reared, and they will disperse and start the process again. The combination of queen and gamergate reproduction results in an increased lifespan of colonies.
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